==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=7-DEC-2009 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HYDROLASE 03-MAY-02 1LNS . COMPND 2 MOLECULE: X-PROLYL DIPEPTIDYL AMINOPETIDASE; . SOURCE 2 ORGANISM_SCIENTIFIC: LACTOCOCCUS LACTIS; . AUTHOR P.RIGOLET,I.MECHIN,M.M.DELAGE,J.F.CHICH . 763 1 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 29616.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 493 64.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 44 5.8 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 118 15.5 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 1 0.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 4 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 5 0.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 80 10.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 69 9.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 143 18.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 9 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 1 1 1 4 1 3 2 2 0 2 1 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 2 1 1 1 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 6 5 2 3 3 2 1 1 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 1 3 1 0 1 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A M 0 0 81 0, 0.0 2,-0.4 0, 0.0 541,-0.0 0.000 360.0 360.0 360.0 162.8 25.3 -8.3 30.5 2 2 A R - 0 0 78 101,-0.1 2,-0.3 2,-0.0 101,-0.2 -0.996 360.0-159.5-137.8 134.7 23.0 -5.5 29.4 3 3 A F - 0 0 1 99,-2.6 2,-0.4 -2,-0.4 503,-0.1 -0.751 1.7-164.9-110.8 159.0 22.7 -3.5 26.2 4 4 A N - 0 0 8 502,-0.4 504,-1.1 -2,-0.3 2,-0.4 -0.972 6.3-163.9-144.2 124.1 21.2 -0.1 25.5 5 5 A H + 0 0 37 -2,-0.4 502,-0.0 98,-0.2 504,-0.0 -0.939 27.4 153.4-118.1 132.8 20.4 1.3 22.0 6 6 A F S S+ 0 0 35 -2,-0.4 523,-0.1 502,-0.2 -1,-0.1 0.349 70.6 70.8-127.2 -4.6 19.7 4.8 21.0 7 7 A S + 0 0 0 118,-0.1 2,-0.6 521,-0.1 -2,-0.0 0.547 65.1 112.2 -92.7 -9.6 20.9 4.6 17.4 8 8 A I - 0 0 41 1,-0.0 2,-0.4 520,-0.0 147,-0.1 -0.544 52.0-158.6 -73.9 114.5 18.1 2.4 16.0 9 9 A V - 0 0 6 -2,-0.6 2,-0.3 145,-0.3 519,-0.1 -0.782 18.7-121.9 -93.5 131.0 16.0 4.4 13.6 10 10 A D - 0 0 137 -2,-0.4 2,-0.3 517,-0.3 -1,-0.0 -0.543 35.6-178.9 -74.4 131.5 12.5 3.2 12.9 11 11 A K - 0 0 21 -2,-0.3 -1,-0.0 143,-0.0 0, 0.0 -0.967 26.7-110.4-134.3 149.9 11.7 2.4 9.3 12 12 A N > - 0 0 110 -2,-0.3 4,-2.5 1,-0.1 5,-0.1 -0.194 37.7-104.9 -68.4 168.4 8.7 1.1 7.3 13 13 A F H > S+ 0 0 33 1,-0.2 4,-2.8 2,-0.2 5,-0.2 0.917 121.1 51.1 -62.3 -45.5 8.7 -2.3 5.8 14 14 A D H > S+ 0 0 126 1,-0.2 4,-1.8 2,-0.2 -1,-0.2 0.880 111.3 49.0 -60.2 -38.7 9.3 -1.0 2.2 15 15 A E H > S+ 0 0 79 2,-0.2 4,-2.3 1,-0.2 -1,-0.2 0.927 110.8 50.8 -66.0 -43.4 12.2 1.0 3.5 16 16 A Q H X S+ 0 0 8 -4,-2.5 4,-2.4 1,-0.2 -2,-0.2 0.920 109.7 49.1 -59.5 -47.6 13.6 -2.1 5.3 17 17 A L H X S+ 0 0 41 -4,-2.8 4,-2.2 1,-0.2 10,-0.2 0.864 111.0 51.0 -61.3 -38.1 13.3 -4.2 2.2 18 18 A A H X S+ 0 0 46 -4,-1.8 4,-1.7 -5,-0.2 -1,-0.2 0.899 111.4 46.1 -67.9 -42.3 15.1 -1.6 0.1 19 19 A E H X S+ 0 0 21 -4,-2.3 4,-0.9 2,-0.2 -2,-0.2 0.892 113.2 49.6 -68.8 -38.1 18.0 -1.2 2.5 20 20 A L H >X>S+ 0 0 0 -4,-2.4 5,-2.5 1,-0.2 3,-0.9 0.916 107.2 55.7 -65.1 -42.2 18.4 -5.0 2.8 21 21 A D H ><5S+ 0 0 84 -4,-2.2 3,-1.6 1,-0.3 -1,-0.2 0.909 103.1 55.4 -56.6 -42.0 18.3 -5.3 -1.0 22 22 A Q H 3<5S+ 0 0 117 -4,-1.7 -1,-0.3 1,-0.3 -2,-0.2 0.769 105.3 52.9 -62.6 -26.3 21.3 -2.9 -1.1 23 23 A L H <<5S- 0 0 14 -4,-0.9 -1,-0.3 -3,-0.9 -2,-0.2 0.508 129.2 -95.9 -87.3 -4.8 23.2 -5.1 1.2 24 24 A G T <<5S+ 0 0 16 -3,-1.6 2,-1.1 -4,-0.5 -3,-0.2 0.474 90.2 123.1 104.7 3.1 22.6 -8.1 -1.1 25 25 A F < + 0 0 2 -5,-2.5 2,-1.3 -6,-0.2 -2,-0.1 -0.722 36.7 180.0 -99.8 83.7 19.6 -9.5 0.7 26 26 A R + 0 0 134 -2,-1.1 2,-0.2 -5,-0.1 -8,-0.1 -0.687 18.8 157.7 -88.3 88.8 17.1 -9.5 -2.1 27 27 A W - 0 0 23 -2,-1.3 2,-0.3 -10,-0.2 -6,-0.1 -0.623 29.4-149.2-110.7 169.5 14.0 -11.0 -0.5 28 28 A S > - 0 0 23 -2,-0.2 3,-1.4 1,-0.0 8,-0.0 -0.938 29.4-113.1-135.2 156.8 10.3 -10.9 -1.1 29 29 A V T 3 S+ 0 0 69 -2,-0.3 85,-0.2 1,-0.3 84,-0.2 0.647 112.2 56.7 -64.9 -17.7 7.3 -11.0 1.2 30 30 A F T 3 S+ 0 0 173 83,-0.1 -1,-0.3 2,-0.1 2,-0.1 0.153 79.6 114.1-103.7 21.1 6.1 -14.4 -0.0 31 31 A W S < S- 0 0 72 -3,-1.4 2,-0.2 1,-0.1 82,-0.1 -0.489 74.9-104.0 -83.5 157.9 9.4 -16.3 0.7 32 32 A D > - 0 0 72 -2,-0.1 4,-2.0 1,-0.1 5,-0.1 -0.574 26.2-123.5 -79.6 150.6 9.4 -19.0 3.3 33 33 A E H > S+ 0 0 30 1,-0.2 4,-2.6 2,-0.2 5,-0.2 0.881 110.9 51.1 -64.7 -37.6 11.0 -18.0 6.6 34 34 A K H > S+ 0 0 21 1,-0.2 4,-2.5 2,-0.2 -1,-0.2 0.854 109.6 52.3 -68.3 -29.7 13.6 -20.9 6.6 35 35 A K H > S+ 0 0 99 2,-0.2 4,-2.6 1,-0.2 -1,-0.2 0.893 109.4 48.9 -69.3 -39.9 14.5 -19.8 3.1 36 36 A I H X S+ 0 0 0 -4,-2.0 4,-2.6 2,-0.2 -2,-0.2 0.935 112.0 48.9 -64.4 -44.7 15.0 -16.3 4.3 37 37 A L H X S+ 0 0 3 -4,-2.6 4,-2.1 2,-0.2 -2,-0.2 0.928 111.8 49.7 -58.6 -47.6 17.1 -17.5 7.2 38 38 A K H X S+ 0 0 80 -4,-2.5 4,-2.6 1,-0.2 -2,-0.2 0.927 112.9 45.7 -58.1 -48.3 19.2 -19.7 4.8 39 39 A D H X S+ 0 0 20 -4,-2.6 4,-2.6 1,-0.2 -1,-0.2 0.898 109.3 56.3 -63.4 -40.4 19.8 -16.8 2.4 40 40 A F H < S+ 0 0 0 -4,-2.6 4,-0.4 1,-0.2 -1,-0.2 0.906 113.6 39.9 -58.0 -43.8 20.7 -14.4 5.3 41 41 A L H >< S+ 0 0 2 -4,-2.1 3,-0.6 2,-0.2 5,-0.3 0.901 115.9 49.4 -73.9 -42.5 23.4 -16.8 6.5 42 42 A I H 3< S+ 0 0 82 -4,-2.6 -2,-0.2 1,-0.2 -1,-0.2 0.849 111.0 49.1 -66.2 -37.1 24.7 -17.8 3.1 43 43 A Q T 3< S+ 0 0 78 -4,-2.6 -1,-0.2 -5,-0.2 -2,-0.2 0.556 103.8 78.3 -79.8 -9.4 25.0 -14.2 1.8 44 44 A S S < S- 0 0 14 -3,-0.6 -3,-0.0 -4,-0.4 -20,-0.0 -0.442 89.6-113.9 -96.6 170.7 26.8 -13.1 5.0 45 45 A P S S+ 0 0 48 0, 0.0 2,-0.1 0, 0.0 -3,-0.1 0.344 80.4 111.8 -84.4 4.9 30.5 -13.6 6.1 46 46 A S S S- 0 0 34 -5,-0.3 2,-0.9 1,-0.1 -2,-0.1 -0.357 74.9-113.4 -79.4 161.1 29.5 -15.9 8.9 47 47 A D > - 0 0 98 1,-0.2 3,-1.5 -2,-0.1 4,-0.5 -0.814 25.4-172.1-100.4 100.0 30.3 -19.6 9.1 48 48 A M G > S+ 0 0 42 -2,-0.9 3,-1.2 1,-0.3 11,-0.4 0.819 79.4 65.4 -57.9 -36.6 27.0 -21.5 8.9 49 49 A T G 3 S+ 0 0 90 1,-0.2 -1,-0.3 10,-0.1 -2,-0.0 0.615 99.8 51.6 -66.4 -13.7 28.7 -24.9 9.6 50 50 A A G < S+ 0 0 74 -3,-1.5 -1,-0.2 8,-0.1 2,-0.2 0.579 89.3 103.2 -97.9 -10.3 29.6 -23.9 13.1 51 51 A L < - 0 0 17 -3,-1.2 8,-2.5 -4,-0.5 2,-0.4 -0.521 59.9-145.2 -82.3 138.1 26.1 -22.7 14.2 52 52 A Q B -A 58 0A 41 6,-0.3 6,-0.3 -2,-0.2 28,-0.1 -0.844 14.3-178.3-100.6 132.2 23.7 -24.6 16.4 53 53 A A - 0 0 0 4,-2.1 2,-0.2 -2,-0.4 5,-0.2 0.840 65.7 -2.6 -97.2 -42.8 20.0 -24.4 15.7 54 54 A T S S- 0 0 18 3,-2.6 -1,-0.2 22,-0.1 38,-0.1 -0.810 83.4 -85.4-139.4-178.6 18.5 -26.6 18.5 55 55 A A S S+ 0 0 70 -2,-0.2 3,-0.1 1,-0.2 37,-0.0 0.682 130.7 27.0 -64.6 -16.5 19.5 -28.8 21.4 56 56 A E S S+ 0 0 161 1,-0.1 2,-0.4 2,-0.1 -1,-0.2 0.587 117.8 54.2-120.2 -17.8 19.8 -31.6 18.9 57 57 A L - 0 0 41 4,-0.0 -3,-2.6 3,-0.0 -4,-2.1 -0.935 60.2-141.3-129.0 146.5 20.6 -30.0 15.5 58 58 A D B > -A 52 0A 53 -2,-0.4 4,-2.3 -6,-0.3 -6,-0.3 -0.337 47.5 -92.9 -85.2 178.9 23.1 -27.6 13.9 59 59 A V H > S+ 0 0 0 -8,-2.5 4,-2.9 -11,-0.4 5,-0.2 0.906 123.9 52.0 -62.0 -45.2 21.9 -25.1 11.3 60 60 A I H > S+ 0 0 54 -12,-0.3 4,-1.8 -9,-0.3 -1,-0.2 0.931 113.2 44.8 -57.1 -47.9 22.6 -27.3 8.3 61 61 A E H > S+ 0 0 91 1,-0.2 4,-1.6 2,-0.2 -1,-0.2 0.886 112.3 53.2 -63.7 -39.7 20.6 -30.2 9.8 62 62 A F H < S+ 0 0 0 -4,-2.3 -2,-0.2 1,-0.2 -1,-0.2 0.919 108.1 49.6 -62.4 -43.9 17.9 -27.8 10.9 63 63 A L H < S+ 0 0 33 -4,-2.9 -1,-0.2 1,-0.2 -2,-0.2 0.859 113.4 46.7 -63.4 -35.3 17.5 -26.5 7.3 64 64 A K H < S+ 0 0 175 -4,-1.8 -1,-0.2 -5,-0.2 -2,-0.2 0.721 103.3 81.4 -78.7 -21.9 17.3 -30.0 6.0 65 65 A S S < S- 0 0 36 -4,-1.6 -3,-0.0 -3,-0.2 0, 0.0 -0.186 83.9-124.1 -78.3 176.1 14.8 -31.1 8.6 66 66 A S S S+ 0 0 121 2,-0.1 -1,-0.1 -2,-0.0 -4,-0.1 0.457 70.4 121.5 -98.3 -5.0 11.1 -30.4 8.5 67 67 A I - 0 0 78 -5,-0.1 2,-0.2 1,-0.1 -2,-0.1 -0.346 61.8-128.6 -59.2 136.7 11.0 -28.6 11.9 68 68 A E - 0 0 99 1,-0.1 2,-0.3 -35,-0.0 -1,-0.1 -0.513 22.3 -99.6 -89.6 158.8 9.7 -25.1 11.4 69 69 A L + 0 0 6 -2,-0.2 2,-0.3 4,-0.1 -1,-0.1 -0.561 43.8 171.9 -78.5 137.3 11.3 -21.9 12.6 70 70 A D > - 0 0 72 -2,-0.3 4,-2.2 1,-0.0 5,-0.1 -0.791 48.7 -92.5-132.6 177.0 10.1 -20.3 15.8 71 71 A W H > S+ 0 0 40 -2,-0.3 4,-2.2 1,-0.2 5,-0.1 0.784 120.9 56.1 -66.6 -27.5 11.4 -17.4 17.9 72 72 A E H > S+ 0 0 48 2,-0.2 4,-2.0 1,-0.2 21,-0.3 0.931 110.7 43.0 -69.2 -45.6 13.5 -19.6 20.2 73 73 A I H > S+ 0 0 9 2,-0.2 4,-2.1 1,-0.2 5,-0.2 0.904 112.1 56.1 -65.8 -40.5 15.4 -21.1 17.3 74 74 A F H X S+ 0 0 5 -4,-2.2 4,-2.9 1,-0.2 -2,-0.2 0.939 111.8 41.7 -55.7 -49.6 15.7 -17.6 15.7 75 75 A W H X S+ 0 0 9 -4,-2.2 4,-2.7 2,-0.2 5,-0.3 0.800 106.6 61.6 -71.2 -28.3 17.3 -16.3 18.9 76 76 A N H X S+ 0 0 4 -4,-2.0 4,-1.0 1,-0.2 -1,-0.2 0.951 115.1 35.9 -59.2 -46.5 19.5 -19.3 19.3 77 77 A I H X S+ 0 0 4 -4,-2.1 4,-2.0 2,-0.2 -2,-0.2 0.902 115.4 54.2 -73.0 -43.7 21.0 -18.4 16.0 78 78 A A H X S+ 0 0 1 -4,-2.9 4,-1.5 1,-0.2 -2,-0.2 0.894 105.8 52.7 -59.9 -42.0 20.9 -14.7 16.4 79 79 A L H <>S+ 0 0 0 -4,-2.7 5,-2.6 1,-0.2 -1,-0.2 0.874 107.9 51.9 -63.5 -35.9 22.8 -14.7 19.7 80 80 A Q H ><5S+ 0 0 21 -4,-1.0 3,-1.8 -5,-0.3 -1,-0.2 0.918 108.2 50.6 -65.6 -41.2 25.6 -16.7 18.1 81 81 A L H 3<5S+ 0 0 9 -4,-2.0 -1,-0.2 1,-0.3 -2,-0.2 0.804 105.7 58.0 -65.3 -26.7 25.8 -14.2 15.3 82 82 A L T 3<5S- 0 0 0 -4,-1.5 47,-3.2 -5,-0.2 48,-0.3 0.259 126.9-102.1 -86.0 11.0 26.0 -11.5 18.0 83 83 A D T < 5S+ 0 0 60 -3,-1.8 2,-0.3 1,-0.3 -3,-0.2 0.681 74.7 141.3 78.3 21.1 29.1 -13.3 19.4 84 84 A F < - 0 0 4 -5,-2.6 -1,-0.3 -6,-0.1 -2,-0.1 -0.759 51.7-124.3 -93.1 144.1 27.5 -15.1 22.3 85 85 A V >>> - 0 0 67 -2,-0.3 4,-2.7 2,-0.0 3,-1.3 -0.775 22.9-133.0 -91.5 114.8 28.7 -18.6 23.0 86 86 A P B 345S+b 90 0B 24 0, 0.0 5,-0.2 0, 0.0 -10,-0.0 -0.356 88.1 17.6 -67.9 147.3 25.9 -21.2 23.1 87 87 A N T 345S+ 0 0 95 3,-2.1 4,-0.1 1,-0.2 -2,-0.0 0.209 132.7 46.6 78.4 -13.9 25.8 -23.6 26.0 88 88 A F T <45S+ 0 0 179 -3,-1.3 -1,-0.2 2,-0.2 3,-0.1 0.696 119.9 27.0-121.8 -54.6 28.1 -21.3 28.0 89 89 A D T <5S+ 0 0 51 -4,-2.7 2,-0.3 1,-0.2 -5,-0.0 0.870 136.0 6.7 -76.6 -36.6 26.9 -17.7 27.8 90 90 A F B - 0 0 119 -2,-0.3 3,-2.6 3,-0.2 7,-0.2 -0.810 34.7-123.4 -96.7 118.8 21.1 -21.9 27.2 92 92 A I T 3 S+ 0 0 16 -2,-0.6 -19,-0.1 1,-0.3 -38,-0.1 -0.353 102.8 30.4 -56.0 132.1 20.1 -23.2 23.8 93 93 A G T 3 S+ 0 0 35 -21,-0.3 -1,-0.3 -20,-0.0 -20,-0.1 0.245 114.5 64.2 98.5 -14.3 16.3 -23.3 23.8 94 94 A K <> + 0 0 103 -3,-2.6 4,-2.0 -22,-0.1 -3,-0.2 -0.023 64.5 115.7-126.5 24.4 16.1 -20.3 26.2 95 95 A A H > S+ 0 0 0 2,-0.2 4,-2.6 1,-0.2 5,-0.2 0.852 70.4 57.1 -64.3 -38.4 17.7 -17.8 23.8 96 96 A F H > S+ 0 0 43 1,-0.2 4,-2.0 2,-0.2 -1,-0.2 0.951 110.3 44.3 -58.2 -50.0 14.6 -15.6 23.5 97 97 A E H > S+ 0 0 79 1,-0.2 4,-2.6 2,-0.2 -1,-0.2 0.893 111.7 52.9 -61.9 -43.0 14.5 -15.1 27.3 98 98 A Y H X S+ 0 0 48 -4,-2.0 4,-1.5 1,-0.2 -1,-0.2 0.927 108.8 50.1 -60.4 -43.3 18.2 -14.5 27.4 99 99 A A H <>S+ 0 0 0 -4,-2.6 5,-2.4 1,-0.2 -1,-0.2 0.898 110.8 50.5 -60.8 -40.1 17.9 -11.8 24.7 100 100 A K H ><5S+ 0 0 110 -4,-2.0 3,-1.6 1,-0.2 -2,-0.2 0.924 109.3 49.4 -64.5 -46.2 15.0 -10.2 26.7 101 101 A N H 3<5S+ 0 0 128 -4,-2.6 -1,-0.2 1,-0.3 -2,-0.2 0.755 107.8 55.1 -66.0 -24.4 17.0 -10.1 29.9 102 102 A S T 3<5S- 0 0 4 -4,-1.5 -99,-2.6 -5,-0.2 -1,-0.3 0.396 113.4-120.6 -87.5 1.8 20.0 -8.5 28.1 103 103 A N T < 5 + 0 0 67 -3,-1.6 -98,-0.2 1,-0.2 -3,-0.2 0.832 53.4 164.8 62.9 34.7 17.7 -5.7 26.9 104 104 A L < - 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0 0 71 -5,-0.2 -6,-2.0 -76,-0.1 -5,-0.4 -0.990 69.8-145.4-133.5 142.2 51.3 29.7 -3.2 332 332 A E E -kL 257 324E 8 -76,-2.2 -74,-2.2 -2,-0.4 2,-0.4 -0.742 11.4-161.8-103.3 151.0 47.6 29.6 -2.2 333 333 A I E -kL 258 323E 7 -10,-2.1 -10,-0.5 -2,-0.3 2,-0.3 -0.988 5.7-150.3-132.3 143.1 45.5 26.5 -1.8 334 334 A K - 0 0 114 -76,-0.5 2,-1.6 -2,-0.4 3,-0.3 -0.809 25.5-116.7-114.9 156.2 41.7 26.1 -1.8 335 335 A A > + 0 0 16 -19,-0.5 3,-1.5 -2,-0.3 5,-0.1 -0.360 54.4 150.0 -84.9 56.8 39.4 23.7 -0.1 336 336 A S T 3 + 0 0 79 -2,-1.6 -1,-0.2 1,-0.3 -136,-0.0 0.685 68.2 54.9 -63.6 -19.5 38.1 22.3 -3.4 337 337 A W T 3 S+ 0 0 31 -3,-0.3 -136,-3.1 -137,-0.1 2,-0.3 0.510 94.2 86.8 -92.5 -6.3 37.5 18.9 -1.9 338 338 A A E < -i 201 0E 7 -3,-1.5 -138,-0.1 -138,-0.3 -135,-0.0 -0.717 60.9-153.3-102.5 147.2 35.3 20.1 1.0 339 339 A N E - 0 0 55 -138,-1.4 -1,-0.1 -2,-0.3 3,-0.1 0.520 47.3-116.8 -90.1 -5.8 31.5 20.7 1.2 340 340 A G E S+ 0 0 2 -139,-0.5 2,-0.4 1,-0.3 -22,-0.1 0.312 73.5 126.3 90.4 -10.4 32.0 23.3 3.9 341 341 A K E - 0 0 46 180,-0.2 -138,-1.9 24,-0.1 2,-0.3 -0.681 37.5-175.9 -86.1 137.3 30.0 21.4 6.6 342 342 A V E -im 203 365E 0 22,-2.9 24,-2.2 -2,-0.4 25,-1.1 -0.967 20.7-173.0-138.7 151.5 31.9 20.9 9.9 343 343 A A E -im 204 367E 0 -140,-1.5 -138,-2.8 -2,-0.3 2,-0.4 -0.982 19.8-140.3-137.8 138.9 31.5 19.2 13.2 344 344 A M E +im 205 368E 1 23,-1.5 25,-2.9 -2,-0.3 2,-0.3 -0.864 23.5 175.9-108.8 140.0 33.9 19.6 16.1 345 345 A T E + m 0 369E 1 -140,-2.0 -138,-0.3 -2,-0.4 2,-0.2 -0.879 21.8 84.3-137.0 167.9 34.9 16.8 18.4 346 346 A G E - m 0 370E 0 23,-0.6 25,-1.9 -2,-0.3 2,-0.3 -0.421 55.9 -96.6 121.5 161.0 37.2 16.0 21.3 347 347 A K E > S- m 0 371E 44 23,-0.2 3,-1.8 -2,-0.2 25,-0.2 -0.733 75.7 -3.2-115.1 165.4 37.3 16.2 25.1 348 348 A S T 3> S+ 0 0 13 23,-1.9 4,-2.4 1,-0.3 26,-0.5 -0.414 136.2 8.8 63.7-122.5 38.7 18.6 27.7 349 349 A Y H 3> S+ 0 0 3 -2,-0.2 4,-1.8 1,-0.2 -1,-0.3 0.856 136.0 52.5 -57.2 -35.2 40.6 21.5 26.0 350 350 A L H <> S+ 0 0 7 -3,-1.8 4,-1.4 2,-0.2 -143,-0.3 0.902 109.4 46.6 -69.7 -41.1 39.3 20.2 22.7 351 351 A G H > S+ 0 0 0 20,-0.4 4,-2.1 1,-0.2 22,-0.2 0.833 108.5 58.6 -67.6 -29.8 35.7 20.2 23.9 352 352 A T H X S+ 0 0 0 -4,-2.4 4,-2.6 1,-0.2 -2,-0.2 0.906 103.5 50.4 -64.4 -43.0 36.4 23.7 25.2 353 353 A M H X S+ 0 0 1 -4,-1.8 4,-3.1 -146,-0.3 -1,-0.2 0.861 107.7 55.0 -65.4 -33.0 37.3 24.9 21.8 354 354 A A H X S+ 0 0 0 -4,-1.4 4,-2.1 2,-0.2 -2,-0.2 0.931 108.9 46.1 -65.8 -45.2 34.1 23.4 20.5 355 355 A Y H X S+ 0 0 0 -4,-2.1 4,-1.6 2,-0.2 -2,-0.2 0.968 116.6 45.8 -59.4 -52.2 32.0 25.4 23.0 356 356 A G H >< S+ 0 0 0 -4,-2.6 3,-0.6 1,-0.2 -2,-0.2 0.941 111.8 49.5 -56.0 -53.4 33.9 28.5 22.2 357 357 A A H ><>S+ 0 0 0 -4,-3.1 3,-1.5 1,-0.2 5,-0.6 0.859 107.3 57.1 -55.4 -37.8 33.8 28.1 18.4 358 358 A A H ><5S+ 0 0 3 -4,-2.1 3,-1.4 1,-0.3 99,-0.5 0.881 98.0 61.1 -63.2 -36.1 30.0 27.5 18.7 359 359 A T T <<5S+ 0 0 2 -4,-1.6 -1,-0.3 -3,-0.6 -2,-0.2 0.443 94.3 64.7 -72.4 2.5 29.6 30.9 20.4 360 360 A T T < 5S- 0 0 38 -3,-1.5 -1,-0.3 -4,-0.2 -2,-0.2 0.613 96.5-136.8 -96.6 -16.2 30.9 32.6 17.3 361 361 A G T < 5 - 0 0 25 -3,-1.4 -3,-0.1 -4,-0.4 -2,-0.1 0.664 33.9-179.5 69.6 17.3 28.1 31.5 15.1 362 362 A V > < - 0 0 23 -5,-0.6 3,-1.2 1,-0.1 -1,-0.2 -0.246 29.9-116.2 -54.8 129.0 30.5 30.7 12.3 363 363 A E T 3 S+ 0 0 147 1,-0.2 -1,-0.1 -22,-0.0 3,-0.1 -0.467 98.3 35.2 -69.9 135.6 28.6 29.5 9.2 364 364 A G T 3 S+ 0 0 7 -2,-0.2 -22,-2.9 1,-0.2 2,-1.5 -0.101 82.7 110.8 112.6 -34.9 29.5 25.9 8.3 365 365 A L E < +m 342 0E 9 -3,-1.2 -1,-0.2 -24,-0.2 -22,-0.2 -0.635 41.6 172.6 -75.3 91.3 29.9 24.7 11.9 366 366 A E E - 0 0 59 -24,-2.2 93,-2.2 -2,-1.5 2,-0.3 0.743 56.5 -10.9 -78.0 -29.1 26.8 22.5 11.9 367 367 A L E -mn 343 459E 0 -25,-1.1 -23,-1.5 91,-0.3 2,-0.4 -0.984 54.3-138.8-168.8 158.1 27.3 20.9 15.2 368 368 A I E -mn 344 460E 0 91,-1.8 93,-2.7 -2,-0.3 2,-0.5 -0.978 2.7-160.3-128.5 142.2 29.6 20.3 18.1 369 369 A L E -mn 345 461E 0 -25,-2.9 2,-0.6 -2,-0.4 -23,-0.6 -0.767 24.3-174.6-118.2 77.3 30.3 17.2 20.2 370 370 A A E -mn 346 462E 0 91,-1.7 93,-2.4 -2,-0.5 2,-0.5 -0.678 2.7-169.8 -83.5 113.8 31.8 18.8 23.3 371 371 A E E -mn 347 463E 2 -25,-1.9 -23,-1.9 -2,-0.6 -20,-0.4 -0.907 68.7 -19.7-110.2 128.6 33.1 16.2 25.7 372 372 A A S S+ 0 0 4 91,-2.6 -1,-0.2 -2,-0.5 -20,-0.2 0.879 97.9 164.6 44.1 47.4 34.2 17.2 29.2 373 373 A G - 0 0 0 90,-0.4 2,-0.5 -3,-0.2 -1,-0.1 -0.471 40.4-117.6 -95.2 166.5 34.6 20.8 28.0 374 374 A I + 0 0 3 -26,-0.5 -18,-0.2 -2,-0.1 3,-0.2 -0.891 21.8 179.4-104.6 125.7 35.0 24.2 29.6 375 375 A S S S+ 0 0 0 -2,-0.5 74,-1.6 1,-0.3 75,-0.8 0.616 82.9 19.0 -97.1 -17.7 32.3 26.8 29.1 376 376 A S B >> -T 448 0G 2 72,-0.2 3,-1.0 73,-0.2 4,-0.6 -0.832 66.5-150.7-155.1 113.4 34.0 29.4 31.3 377 377 A W H >> S+ 0 0 2 70,-2.1 4,-1.3 -2,-0.3 3,-1.1 0.786 91.2 70.4 -55.5 -31.9 37.6 29.2 32.3 378 378 A Y H >> S+ 0 0 0 66,-0.3 4,-1.5 1,-0.3 3,-0.5 0.916 98.1 49.8 -52.9 -45.9 37.0 31.0 35.6 379 379 A N H <4 S+ 0 0 24 -3,-1.0 -1,-0.3 1,-0.2 -2,-0.2 0.692 102.9 61.7 -69.3 -17.4 35.2 27.9 37.0 380 380 A Y H << S+ 0 0 14 -3,-1.1 -1,-0.2 -4,-0.6 -2,-0.2 0.857 122.5 19.0 -77.0 -33.1 38.0 25.6 35.9 381 381 A Y H << S+ 0 0 1 -4,-1.3 7,-1.3 -3,-0.5 2,-0.3 0.376 129.5 37.7-117.0 3.0 40.6 27.3 38.1 382 382 A R B < -U 387 0H 0 -4,-1.5 2,-0.2 5,-0.2 5,-0.2 -0.920 53.5-163.3-146.5 169.3 38.3 29.2 40.6 383 383 A E S S- 0 0 5 3,-2.5 3,-0.5 -2,-0.3 304,-0.1 -0.802 71.6 -24.2-162.8 114.7 35.1 28.8 42.5 384 384 A N S S- 0 0 46 1,-0.3 303,-2.0 -2,-0.2 306,-0.2 0.906 128.4 -39.2 46.1 61.9 33.1 31.6 44.2 385 385 A G S S+ 0 0 0 53,-0.2 2,-0.4 1,-0.2 -1,-0.3 0.703 120.4 110.7 67.3 20.9 35.9 34.1 44.7 386 386 A L S S- 0 0 9 -3,-0.5 -3,-2.5 52,-0.2 2,-0.7 -0.965 74.8-115.4-132.5 146.2 38.3 31.3 45.6 387 387 A V B +U 382 0H 8 -2,-0.4 2,-0.5 -5,-0.2 350,-0.4 -0.678 40.4 176.1 -75.6 114.6 41.3 29.7 44.0 388 388 A R - 0 0 6 -7,-1.3 348,-0.2 -2,-0.7 3,-0.1 -0.897 20.8-141.7-131.6 102.6 40.2 26.2 43.4 389 389 A S - 0 0 0 346,-2.7 3,-0.1 -2,-0.5 -8,-0.1 -0.236 33.8 -94.1 -61.9 144.4 42.6 23.8 41.6 390 390 A P > - 0 0 1 0, 0.0 3,-2.7 0, 0.0 80,-0.2 -0.297 60.6 -80.8 -57.9 144.0 41.3 21.3 39.1 391 391 A G T 3 S+ 0 0 8 78,-2.0 344,-0.1 79,-0.3 78,-0.0 -0.184 117.3 10.3 -48.7 131.5 40.6 18.0 40.7 392 392 A G T 3 S+ 0 0 34 -3,-0.1 -1,-0.3 342,-0.1 78,-0.1 0.345 114.6 84.2 80.7 -7.0 43.8 16.0 41.0 393 393 A F X - 0 0 109 -3,-2.7 3,-1.4 341,-0.2 341,-0.1 -0.540 57.4-175.5-131.0 72.4 46.0 18.9 40.0 394 394 A P T 3 S+ 0 0 36 0, 0.0 3,-0.1 0, 0.0 341,-0.1 -0.349 72.3 26.3 -65.9 144.4 46.9 21.3 42.9 395 395 A G T 3 S+ 0 0 0 1,-0.3 -171,-3.1 340,-0.2 2,-0.2 0.413 89.7 131.6 85.9 -1.8 48.9 24.4 42.0 396 396 A E < + 0 0 21 -3,-1.4 -1,-0.3 -173,-0.2 2,-0.3 -0.581 21.7 152.9 -87.7 148.3 47.6 24.5 38.4 397 397 A D >> - 0 0 0 -2,-0.2 4,-1.8 -3,-0.1 3,-0.7 -0.911 60.3 -77.3-155.9 178.2 46.3 27.7 36.8 398 398 A L H 3> S+ 0 0 0 -2,-0.3 4,-2.1 1,-0.2 5,-0.1 0.810 126.4 54.3 -57.4 -32.8 45.9 29.2 33.3 399 399 A D H 3> S+ 0 0 5 2,-0.2 4,-1.9 1,-0.2 -1,-0.2 0.837 105.5 52.7 -72.5 -30.6 49.6 30.2 33.1 400 400 A V H <> S+ 0 0 0 -3,-0.7 4,-1.6 2,-0.2 -2,-0.2 0.904 111.8 46.5 -68.7 -39.7 50.8 26.6 33.9 401 401 A L H X S+ 0 0 4 -4,-1.8 4,-2.6 2,-0.2 -2,-0.2 0.877 108.5 56.0 -68.5 -37.3 48.6 25.4 31.0 402 402 A A H X S+ 0 0 0 -4,-2.1 4,-0.8 1,-0.2 -2,-0.2 0.915 107.7 48.4 -61.1 -42.8 49.9 28.1 28.8 403 403 A A H < S+ 0 0 2 -4,-1.9 3,-0.3 1,-0.2 -1,-0.2 0.879 111.6 50.9 -65.6 -35.6 53.5 26.9 29.4 404 404 A L H < S+ 0 0 21 -4,-1.6 -189,-2.2 1,-0.2 -2,-0.2 0.911 118.1 36.4 -67.9 -41.4 52.4 23.3 28.7 405 405 A T H < S+ 0 0 0 -4,-2.6 2,-2.0 -191,-0.2 -1,-0.2 0.312 81.8 102.4 -99.9 8.2 50.7 24.1 25.4 406 406 A Y >< + 0 0 1 -4,-0.8 3,-1.2 -3,-0.3 -114,-0.2 -0.567 45.9 165.6 -85.8 74.5 53.1 26.8 24.1 407 407 A S G > + 0 0 0 -2,-2.0 3,-1.6 -116,-0.3 4,-0.3 0.459 52.5 87.0 -75.9 -2.9 54.5 24.1 21.8 408 408 A R G > S+ 0 0 5 -117,-1.0 3,-1.6 1,-0.3 7,-0.3 0.883 84.8 60.4 -61.9 -34.8 56.3 26.7 19.6 409 409 A N G < S+ 0 0 19 -3,-1.2 -1,-0.3 1,-0.3 -2,-0.2 0.545 83.6 78.5 -70.3 -5.7 59.1 26.4 22.1 410 410 A L G < S+ 0 0 50 -3,-1.6 2,-0.6 -197,-0.1 -1,-0.3 0.695 84.3 72.5 -75.6 -16.0 59.5 22.6 21.4 411 411 A D S <> S- 0 0 48 -3,-1.6 4,-2.8 -4,-0.3 3,-0.5 -0.884 77.2-148.6 -99.6 123.3 61.4 23.6 18.3 412 412 A G H > S+ 0 0 54 -2,-0.6 4,-2.3 1,-0.2 -1,-0.2 0.900 98.4 53.9 -55.3 -44.2 64.8 24.9 19.0 413 413 A A H > S+ 0 0 79 1,-0.2 4,-1.1 2,-0.2 -1,-0.2 0.839 112.4 44.5 -60.1 -35.0 64.8 27.3 16.0 414 414 A D H > S+ 0 0 9 -3,-0.5 4,-2.5 -6,-0.3 5,-0.3 0.892 107.6 57.8 -76.5 -42.2 61.5 28.8 17.3 415 415 A F H X S+ 0 0 95 -4,-2.8 4,-1.8 -7,-0.3 5,-0.3 0.926 104.5 53.8 -51.6 -48.4 62.7 29.0 20.9 416 416 A L H < S+ 0 0 153 -4,-2.3 -1,-0.2 1,-0.2 4,-0.2 0.898 116.1 37.3 -54.2 -45.5 65.6 31.2 19.6 417 417 A K H < S+ 0 0 140 -4,-1.1 -1,-0.2 -5,-0.2 -2,-0.2 0.728 130.9 20.6 -84.1 -23.6 63.3 33.6 17.9 418 418 A G H X S+ 0 0 3 -4,-2.5 4,-1.6 2,-0.1 -3,-0.2 0.354 87.8 96.8-133.8 13.4 60.3 33.9 20.2 419 419 A N H X S+ 0 0 26 -4,-1.8 4,-2.6 -5,-0.3 5,-0.3 0.921 86.5 53.0 -71.1 -39.9 61.2 32.7 23.8 420 420 A A H > S+ 0 0 78 -5,-0.3 4,-2.0 1,-0.2 -1,-0.2 0.944 111.5 44.8 -58.7 -50.5 61.8 36.3 25.0 421 421 A E H > S+ 0 0 62 1,-0.2 4,-2.1 2,-0.2 -1,-0.2 0.846 112.4 55.6 -62.4 -34.7 58.4 37.5 23.8 422 422 A Y H X S+ 0 0 15 -4,-1.6 4,-2.1 2,-0.2 -2,-0.2 0.932 108.0 44.4 -64.8 -50.0 56.8 34.4 25.3 423 423 A E H X S+ 0 0 112 -4,-2.6 4,-2.3 2,-0.2 5,-0.2 0.889 111.8 55.5 -63.7 -37.7 58.1 34.8 28.8 424 424 A K H X S+ 0 0 135 -4,-2.0 4,-1.5 -5,-0.3 -1,-0.2 0.935 112.9 40.8 -59.4 -47.8 57.2 38.5 28.7 425 425 A R H X S+ 0 0 36 -4,-2.1 4,-1.7 1,-0.2 -1,-0.2 0.781 112.1 54.9 -74.0 -26.1 53.6 37.6 27.8 426 426 A L H X S+ 0 0 27 -4,-2.1 4,-2.7 2,-0.2 -1,-0.2 0.876 107.2 50.9 -73.3 -37.0 53.4 34.7 30.3 427 427 A A H X S+ 0 0 63 -4,-2.3 4,-1.9 1,-0.2 -2,-0.2 0.903 109.4 51.1 -64.9 -40.8 54.4 37.0 33.1 428 428 A E H X S+ 0 0 129 -4,-1.5 4,-1.8 -5,-0.2 -1,-0.2 0.886 111.1 48.4 -63.5 -38.8 51.7 39.4 32.1 429 429 A M H X S+ 0 0 8 -4,-1.7 4,-1.6 2,-0.2 -2,-0.2 0.945 107.9 54.4 -66.4 -47.0 49.2 36.5 32.1 430 430 A T H X S+ 0 0 55 -4,-2.7 4,-0.6 1,-0.2 5,-0.3 0.865 110.1 46.6 -54.8 -40.5 50.4 35.3 35.5 431 431 A A H >< S+ 0 0 81 -4,-1.9 3,-0.8 1,-0.2 -1,-0.2 0.879 108.8 55.3 -71.0 -36.7 49.8 38.8 37.0 432 432 A A H 3< S+ 0 0 35 -4,-1.8 -1,-0.2 1,-0.2 -2,-0.2 0.777 94.9 69.1 -66.5 -25.1 46.4 39.1 35.3 433 433 A L H 3< S- 0 0 0 -4,-1.6 -1,-0.2 -3,-0.1 -2,-0.2 0.820 85.1-157.9 -62.5 -33.4 45.3 35.8 37.0 434 434 A D - 0 0 31 -2,-0.1 4,-2.2 -203,-0.1 5,-0.1 -0.885 53.4 -78.0-154.9-175.1 37.9 39.9 36.6 442 442 A Q H > S+ 0 0 52 -2,-0.3 4,-1.7 1,-0.2 5,-0.1 0.864 123.3 57.7 -62.0 -39.1 38.1 41.2 33.0 443 443 A F H 4 S+ 0 0 36 1,-0.2 -1,-0.2 -11,-0.2 3,-0.2 0.912 112.1 41.5 -57.8 -44.7 40.8 38.7 32.1 444 444 A W H >4 S+ 0 0 0 1,-0.2 3,-2.0 -12,-0.2 -66,-0.3 0.870 108.7 60.5 -71.0 -36.9 38.5 35.9 33.1 445 445 A H H >< S+ 0 0 25 -4,-2.2 3,-1.2 1,-0.3 -1,-0.2 0.800 97.8 58.4 -61.5 -30.1 35.5 37.6 31.4 446 446 A D T 3< S+ 0 0 39 -4,-1.7 -142,-0.5 1,-0.3 -1,-0.3 0.517 106.9 50.0 -77.4 -4.3 37.3 37.4 28.1 447 447 A R T < S+ 0 0 14 -3,-2.0 -70,-2.1 -144,-0.1 2,-0.6 -0.102 84.9 103.8-122.8 31.9 37.4 33.7 28.5 448 448 A N B X +T 376 0G 17 -3,-1.2 3,-0.6 -72,-0.2 -72,-0.2 -0.934 40.3 177.7-118.5 104.4 33.7 33.2 29.4 449 449 A Y G > + 0 0 0 -74,-1.6 3,-2.1 -2,-0.6 -73,-0.2 0.640 68.1 83.2 -80.6 -16.9 31.8 31.8 26.4 450 450 A L G > S+ 0 0 33 -75,-0.8 3,-1.1 1,-0.3 -1,-0.2 0.794 83.3 61.2 -57.4 -29.5 28.5 31.6 28.4 451 451 A I G < S+ 0 0 54 -3,-0.6 -1,-0.3 1,-0.2 -2,-0.1 0.518 105.8 47.5 -77.9 -2.3 27.9 35.3 27.6 452 452 A N G X S+ 0 0 2 -3,-2.1 3,-1.8 1,-0.1 -1,-0.2 0.175 71.5 110.2-122.2 16.9 27.8 34.5 23.8 453 453 A T G X + 0 0 10 -3,-1.1 3,-1.2 1,-0.3 -1,-0.1 0.639 66.7 73.9 -67.2 -12.7 25.5 31.5 23.7 454 454 A D G 3 S+ 0 0 95 1,-0.2 -1,-0.3 -3,-0.1 -2,-0.1 0.651 93.0 55.8 -74.0 -12.7 22.9 33.7 22.0 455 455 A K G < S+ 0 0 87 -3,-1.8 2,-0.5 -96,-0.1 -1,-0.2 0.407 79.5 118.1 -98.0 1.8 25.1 33.5 18.9 456 456 A V < + 0 0 0 -3,-1.2 -97,-0.1 1,-0.2 -98,-0.1 -0.576 23.6 157.7 -74.8 120.2 25.0 29.6 18.9 457 457 A K + 0 0 138 -2,-0.5 -1,-0.2 -99,-0.5 2,-0.1 0.608 46.0 93.3-113.6 -22.6 23.4 28.2 15.7 458 458 A A S S- 0 0 5 -100,-0.3 -91,-0.3 1,-0.1 -92,-0.1 -0.406 86.9-101.0 -74.2 147.3 25.0 24.8 15.7 459 459 A D E -n 367 0E 15 -93,-2.2 -91,-1.8 -2,-0.1 2,-0.6 -0.397 41.1-136.4 -61.7 146.9 23.2 21.8 17.2 460 460 A V E +no 368 489E 0 28,-2.4 30,-2.4 -93,-0.2 2,-0.5 -0.930 24.3 179.7-118.1 116.1 24.8 21.1 20.6 461 461 A L E -no 369 490E 0 -93,-2.7 -91,-1.7 -2,-0.6 2,-0.4 -0.962 13.5-166.1-112.1 124.4 25.7 17.6 21.8 462 462 A I E -no 370 491E 2 28,-3.1 30,-3.1 -2,-0.5 2,-0.4 -0.907 10.1-173.4-114.1 138.7 27.3 17.5 25.2 463 463 A V E +no 371 492E 1 -93,-2.4 -91,-2.6 -2,-0.4 -90,-0.4 -0.996 16.7 159.9-129.1 134.1 29.1 14.6 26.9 464 464 A H E - 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