==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=1-JUL-2012 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER FLUORESCENT PROTEIN 06-MAY-09 2WHS . COMPND 2 MOLECULE: LARGE STOKES SHIFT FLUORESCENT PROTEIN; . SOURCE 2 ORGANISM_SCIENTIFIC: MONTIPORA SP. 20; . AUTHOR S.VIOLOT,P.CARPENTIER,L.BLANCHOIN,D.BOURGEOIS . 883 8 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 38088.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 622 70.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 52 5.9 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 409 46.3 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 8 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 8 0.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 12 1.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 62 7.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 69 7.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 24 2.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 7 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 4 0 0 0 0 0 0 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 0 4 0 0 8 0 4 4 8 3 13 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 0 0 0 0 0 0 0 0 0 0 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 1 A V 0 0 79 0, 0.0 2,-1.7 0, 0.0 3,-0.4 0.000 360.0 360.0 360.0 21.1 -41.3 -3.7 -12.1 2 2 A S + 0 0 113 1,-0.3 77,-0.0 2,-0.2 76,-0.0 -0.344 360.0 53.1 87.7 -72.2 -42.8 -2.4 -8.8 3 3 A V S S+ 0 0 54 -2,-1.7 2,-0.6 75,-0.1 -1,-0.3 0.685 104.1 76.1 -72.3 -17.6 -40.2 -0.4 -6.9 4 4 A I S S- 0 0 6 -3,-0.4 -2,-0.2 77,-0.1 -1,-0.1 -0.857 73.0-157.4-106.3 114.6 -39.8 1.4 -10.2 5 5 A A - 0 0 31 -2,-0.6 29,-0.2 2,-0.2 3,-0.1 -0.290 31.6-107.2 -83.8 168.3 -42.5 4.0 -11.1 6 6 A K S S+ 0 0 173 1,-0.2 2,-0.4 26,-0.1 28,-0.3 0.786 113.0 40.1 -66.5 -28.5 -43.4 5.3 -14.6 7 7 A Q S S+ 0 0 109 25,-0.2 2,-0.3 26,-0.1 -1,-0.2 -0.988 76.8 166.2-129.8 120.9 -41.8 8.6 -13.6 8 8 A M E -A 31 0A 2 23,-2.0 23,-2.1 -2,-0.4 2,-0.2 -0.906 19.3-153.7-129.8 152.2 -38.6 8.6 -11.6 9 9 A T E -Ab 30 111A 66 101,-2.5 103,-2.4 -2,-0.3 2,-0.3 -0.648 13.8-157.4-111.1 179.2 -35.9 11.2 -10.7 10 10 A Y E -Ab 29 112A 4 19,-2.3 19,-2.7 101,-0.3 2,-0.4 -0.991 15.8-148.6-157.2 155.2 -32.2 10.7 -9.8 11 11 A K E -Ab 28 113A 38 101,-2.8 103,-2.6 -2,-0.3 2,-0.5 -0.989 18.8-167.3-117.4 126.6 -29.2 12.2 -8.1 12 12 A V E -Ab 27 114A 1 15,-2.4 15,-2.3 -2,-0.4 2,-0.5 -0.971 6.4-167.9-116.6 131.2 -25.7 11.5 -9.4 13 13 A Y E -Ab 26 115A 112 101,-3.1 103,-3.6 -2,-0.5 2,-0.4 -0.966 14.0-179.1-110.0 123.6 -22.5 12.3 -7.6 14 14 A M E +Ab 25 116A 4 11,-3.6 11,-3.0 -2,-0.5 2,-0.3 -0.960 7.4 163.1-123.0 139.2 -19.4 12.1 -9.7 15 15 A S E +Ab 24 117A 26 101,-2.3 103,-1.7 -2,-0.4 2,-0.3 -0.998 15.5 110.1-143.1 152.3 -15.8 12.6 -8.7 16 16 A G E -Ab 23 118A 8 7,-2.3 7,-2.4 -2,-0.3 2,-0.4 -0.996 50.6 -86.7 175.9-161.4 -12.5 11.8 -10.1 17 17 A T E -Ab 22 119A 37 101,-2.0 103,-2.7 -2,-0.3 2,-0.5 -0.995 21.9-169.7-133.8 128.9 -9.2 12.8 -11.8 18 18 A V E > S-Ab 21 120A 0 3,-2.3 3,-2.1 -2,-0.4 103,-0.2 -0.972 80.9 -21.3-120.1 120.1 -8.4 13.2 -15.5 19 19 A N T 3 S- 0 0 31 101,-3.5 652,-0.4 -2,-0.5 -1,-0.2 0.929 131.1 -48.7 49.1 48.2 -4.7 13.7 -16.4 20 20 A G T 3 S+ 0 0 12 100,-0.4 649,-1.9 1,-0.2 2,-0.7 0.390 110.8 125.6 77.6 -6.3 -4.1 14.7 -12.8 21 21 A H E < -A 18 0A 59 -3,-2.1 -3,-2.3 647,-0.2 2,-0.3 -0.808 49.5-149.2 -92.1 116.5 -7.0 17.2 -12.7 22 22 A Y E +A 17 0A 27 -2,-0.7 2,-0.3 -5,-0.2 -5,-0.2 -0.601 26.7 158.5 -80.9 138.7 -9.4 16.6 -9.8 23 23 A F E -A 16 0A 0 -7,-2.4 -7,-2.3 -2,-0.3 2,-0.4 -0.990 28.2-147.7-152.3 162.2 -13.1 17.4 -10.1 24 24 A E E -AC 15 46A 64 22,-2.2 21,-1.6 -2,-0.3 22,-1.5 -0.986 11.8-168.5-130.9 145.2 -16.5 16.8 -8.7 25 25 A V E -AC 14 44A 3 -11,-3.0 -11,-3.6 -2,-0.4 2,-0.4 -0.988 1.8-167.7-129.2 143.7 -19.9 16.8 -10.5 26 26 A E E -AC 13 43A 65 17,-2.5 17,-2.5 -2,-0.4 2,-0.3 -0.988 11.6-175.9-125.7 139.4 -23.4 16.7 -9.1 27 27 A G E -AC 12 42A 1 -15,-2.3 -15,-2.4 -2,-0.4 2,-0.5 -0.989 22.2-155.0-137.9 151.2 -26.6 16.2 -11.2 28 28 A D E +AC 11 41A 91 13,-2.1 13,-2.8 -2,-0.3 2,-0.2 -0.995 29.7 175.7-117.5 121.5 -30.4 16.0 -10.9 29 29 A G E -AC 10 40A 5 -19,-2.7 -19,-2.3 -2,-0.5 2,-0.3 -0.740 14.5-173.3-124.2 171.1 -32.1 13.9 -13.5 30 30 A K E +AC 9 39A 120 9,-2.7 9,-2.6 -21,-0.3 2,-0.3 -0.973 17.2 136.3-161.3 156.2 -35.6 12.7 -14.4 31 31 A G E -AC 8 38A 7 -23,-2.1 -23,-2.0 -2,-0.3 7,-0.2 -0.987 53.0 -85.7 177.8-175.9 -37.1 10.3 -16.9 32 32 A K E >>> - C 0 37A 77 5,-2.5 3,-2.3 -2,-0.3 4,-1.2 -0.882 31.5-165.0-122.1 91.9 -39.5 7.6 -17.8 33 33 A P T 345S+ 0 0 0 0, 0.0 3,-0.1 0, 0.0 -1,-0.1 0.770 87.9 48.8 -55.6 -31.5 -37.8 4.3 -17.0 34 34 A Y T 345S+ 0 0 78 -28,-0.3 -28,-0.1 1,-0.2 32,-0.1 0.496 114.9 46.8 -86.4 -4.7 -40.2 2.2 -19.1 35 35 A E T <45S- 0 0 98 -3,-2.3 33,-0.2 -29,-0.2 -1,-0.2 0.489 105.0-125.9-109.8 -8.2 -39.9 4.5 -22.1 36 36 A G T <5 + 0 0 0 -4,-1.2 175,-1.9 1,-0.3 2,-0.4 0.656 64.8 132.5 75.3 17.2 -36.1 4.8 -22.1 37 37 A E E < +CD 32 210A 93 -5,-1.0 -5,-2.5 173,-0.2 2,-0.3 -0.869 25.8 164.4-108.0 140.7 -36.1 8.6 -21.9 38 38 A Q E -CD 31 209A 14 171,-1.7 171,-2.3 -2,-0.4 2,-0.3 -0.991 17.0-173.8-154.3 144.4 -33.9 10.5 -19.5 39 39 A T E -CD 30 208A 39 -9,-2.6 -9,-2.7 -2,-0.3 2,-0.3 -0.981 8.4-174.0-139.0 147.7 -32.7 14.0 -18.9 40 40 A V E -CD 29 207A 3 167,-2.7 167,-2.6 -2,-0.3 2,-0.5 -0.998 15.6-147.4-143.0 143.1 -30.1 15.3 -16.3 41 41 A K E -CD 28 206A 124 -13,-2.8 -13,-2.1 -2,-0.3 2,-0.3 -0.950 24.5-160.4-111.7 131.1 -28.9 18.7 -15.3 42 42 A L E -CD 27 205A 3 163,-3.7 163,-1.7 -2,-0.5 2,-0.4 -0.832 15.4-167.9-119.4 148.3 -25.2 18.9 -14.1 43 43 A T E -CD 26 204A 72 -17,-2.5 -17,-2.5 -2,-0.3 2,-0.8 -0.998 24.4-133.2-131.0 128.5 -23.1 21.2 -12.1 44 44 A V E +C 25 0A 18 159,-2.4 -19,-0.3 -2,-0.4 4,-0.1 -0.733 30.2 179.2 -73.8 109.5 -19.3 21.1 -11.8 45 45 A T E + 0 0 66 -21,-1.6 2,-0.3 -2,-0.8 -20,-0.2 0.706 61.5 18.6 -93.7 -23.4 -19.1 21.4 -8.0 46 46 A K E S+C 24 0A 106 -22,-1.5 -22,-2.2 1,-0.0 -1,-0.2 -0.989 119.4 20.3-145.5 142.8 -15.2 21.3 -7.5 47 47 A G S S- 0 0 27 -2,-0.3 -2,-0.1 1,-0.2 -1,-0.0 0.655 88.2-171.5 78.9 18.9 -12.3 21.7 -10.0 48 48 A G + 0 0 39 -4,-0.1 -1,-0.2 2,-0.1 2,-0.1 -0.630 56.2 49.5 -98.1 160.7 -14.1 23.7 -12.7 49 49 A P S S- 0 0 70 0, 0.0 153,-0.1 0, 0.0 154,-0.1 0.555 88.8-134.0 -65.2 145.6 -13.8 24.8 -15.4 50 50 A L - 0 0 7 151,-0.4 4,-0.1 152,-0.2 -2,-0.1 -0.498 2.9-144.9 -69.7 137.8 -12.6 21.4 -16.4 51 51 A P S S+ 0 0 49 0, 0.0 2,-0.3 0, 0.0 -1,-0.1 0.194 79.2 67.2 -83.2 13.3 -9.4 21.5 -18.6 52 52 A F S S- 0 0 4 79,-0.1 78,-0.2 80,-0.1 2,-0.2 -0.870 99.7 -74.3-129.9 164.9 -10.7 18.5 -20.6 53 53 A A > - 0 0 4 76,-1.7 3,-1.4 -2,-0.3 4,-0.4 -0.356 32.6-141.4 -58.0 124.1 -13.6 17.6 -23.0 54 54 A W G >> S+ 0 0 9 1,-0.2 3,-2.6 -2,-0.2 4,-1.1 0.867 92.6 76.1 -50.3 -37.1 -17.0 17.3 -21.2 55 55 A D G 34 S+ 0 0 11 1,-0.3 -1,-0.2 2,-0.2 82,-0.1 0.691 80.9 63.4 -57.6 -29.2 -17.8 14.4 -23.4 56 56 A I G <4 S+ 0 0 0 -3,-1.4 -1,-0.3 1,-0.2 -2,-0.2 0.785 108.9 44.6 -67.4 -20.7 -15.7 11.9 -21.7 57 57 A L T X4 S+ 0 0 0 -3,-2.6 3,-2.6 -4,-0.4 4,-0.4 0.818 93.0 80.9 -86.3 -32.9 -17.9 12.4 -18.6 58 58 A S G >< S+ 0 0 2 -4,-1.1 3,-1.5 1,-0.3 -2,-0.1 0.817 88.0 52.1 -54.6 -41.9 -21.3 12.3 -20.3 59 59 A P G 3 S+ 0 0 20 0, 0.0 -1,-0.3 0, 0.0 -2,-0.1 0.504 104.1 57.3 -77.0 2.8 -21.7 8.5 -20.5 60 60 A Q G < 0 0 4 -3,-2.6 -2,-0.2 1,-0.1 -3,-0.1 0.397 360.0 360.0-100.5 -0.2 -21.0 7.9 -16.8 61 61 A L < 0 0 9 -3,-1.5 -1,-0.1 -4,-0.4 -3,-0.1 0.620 360.0 360.0-102.5 360.0 -23.8 10.2 -15.9 62 ! 0 0 0 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 360.0 0.0 0.0 0.0 63 65 A S > 0 0 15 0, 0.0 3,-1.7 0, 0.0 -2,-0.1 0.000 360.0 360.0 360.0 69.0 -28.2 5.9 -15.7 64 66 A I G > + 0 0 6 1,-0.3 3,-0.9 2,-0.1 14,-0.2 0.622 360.0 72.0 -66.3 -27.2 -29.2 2.4 -16.8 65 67 A P G 3 S+ 0 0 0 0, 0.0 -1,-0.3 0, 0.0 -29,-0.1 0.712 86.6 73.0 -61.0 -14.1 -32.6 1.7 -15.2 66 68 A F G < S+ 0 0 0 -3,-1.7 2,-0.5 -30,-0.1 145,-0.4 -0.090 72.0 118.8 -86.5 35.4 -34.0 4.2 -17.7 67 69 A T < - 0 0 1 -3,-0.9 2,-0.7 143,-0.1 145,-0.2 -0.909 67.7-125.1-103.0 127.6 -33.6 1.7 -20.5 68 70 A K B -e 212 0A 85 143,-2.2 145,-2.4 -2,-0.5 -34,-0.1 -0.600 31.0-171.7 -69.9 110.7 -36.8 0.5 -22.4 69 71 A Y - 0 0 29 -2,-0.7 2,-0.1 143,-0.2 6,-0.1 -0.887 20.4-129.0-102.7 132.1 -36.9 -3.2 -22.2 70 72 A P > - 0 0 51 0, 0.0 3,-1.7 0, 0.0 144,-0.0 -0.460 34.3-104.3 -69.9 158.9 -39.5 -5.2 -24.2 71 73 A E T 3 S+ 0 0 207 1,-0.3 4,-0.1 -2,-0.1 -2,-0.0 0.732 115.9 58.5 -54.2 -31.5 -41.5 -7.7 -22.1 72 74 A D T 3 S+ 0 0 108 1,-0.1 -1,-0.3 2,-0.1 -3,-0.0 0.515 94.1 69.8 -87.7 -3.2 -39.6 -10.8 -23.4 73 75 A I S < S- 0 0 8 -3,-1.7 -1,-0.1 111,-0.1 140,-0.1 -0.942 88.8-122.6-111.4 115.7 -36.1 -9.7 -22.3 74 76 A P - 0 0 34 0, 0.0 2,-1.1 0, 0.0 3,-0.1 -0.273 21.1-130.2 -52.9 134.7 -35.6 -9.7 -18.5 75 77 A D > - 0 0 15 1,-0.2 4,-2.7 -4,-0.1 5,-0.2 -0.768 23.8-173.8 -95.4 93.1 -34.6 -6.2 -17.3 76 78 A Y H > S+ 0 0 3 -2,-1.1 4,-1.0 1,-0.2 -1,-0.2 0.849 80.9 47.6 -52.2 -44.1 -31.5 -6.9 -15.2 77 79 A F H >4 S+ 0 0 0 2,-0.2 3,-0.6 1,-0.2 4,-0.4 0.939 113.0 44.3 -72.2 -49.9 -31.2 -3.3 -14.0 78 80 A K H >4 S+ 0 0 2 1,-0.2 3,-1.6 2,-0.2 -2,-0.2 0.939 110.7 56.8 -60.6 -45.1 -34.8 -2.7 -13.0 79 81 A Q H 3< S+ 0 0 26 -4,-2.7 -1,-0.2 1,-0.3 -2,-0.2 0.780 93.4 69.3 -53.6 -31.5 -34.9 -6.1 -11.2 80 82 A S T - 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